Leptonetela Kratochvíl, 1978 that is endemic to karst systems in Eurasia using DNA barcoding. We analyzed 624 specimens using one mitochondrial gene fragment (COI). The results show that DNA barcoding is an efficient and rapid species identification method in this genus. DNA barcoding gap and automatic barcode gap discovery (ABGD) analyses indicated the existence of 90 species, a result consistent with previous taxonomic hypotheses, and supported the existence of extreme male pedipalpal tibial spine and median apophysis polymorphism in Leptonetela species, with direct implications for the taxonomy of the group and its diversity. Based on the molecular and morphological evidence, we delimit and diagnose 90 Leptonetela species, including the type species Leptonetela kanellisi. Forty-six of them are previously undescribed. The female of Leptonetela zhaiLeptonetela tianxinensis  comb. nov. is transferred from the genus Leptoneta Simon, 1872;the genus Guineta Lin & Li, 2010 syn. nov. is a junior synonym of Leptonetela; Leptonetela gigachela comb. nov. is transferred from Guineta. The genus Sinoneta Lin & Li, 2010 syn. nov. is a junior synonym of Leptonetela; Leptonetela notabilis comb. nov. and Leptonetela sexdigiti comb. nov. are transferred from Sinoneta; Leptonetela sanchahe Wang & Li nom. nov. is proposed as a replacement name for Sinoneta palmata because Leptonetela palmata is preoccupied.Extreme environments, such as subterranean habitats, are suspected to be responsible for morphologically inseparable cryptic or sibling species and can bias biodiversity assessment. A DNA barcode is a short, standardized DNA sequence used for taxonomic purposes and has the potential to lessen the challenges presented by a biotic inventory. Here, we investigate the diversity of the genus  Subterranean ecosystems, such as caves and cracks, are evident mainly in karst areas, which represent nearly 4% of the rocky outcrops of the world. These environments are marked by permanent darkness, a lack of diurnal and annual rhythms, and extremely scarce food sources . Many stDNA barcoding relies on the use of a standardized DNA region as a tag for accurate and rapid species identification  and has Leptonetela is discontinuously distributed in the South China karst and the Balkan Peninsula, a karstic region in Europe. The genus has 54 catalogued species . Rapid and accurate identification within this genus is difficult due to congeneric species sharing similar morphological traits, a lack of obvious morphological differences between closely related species and some species only differ in one or a few quantitative differences, such as the location, length ratio or thickness of the male pedipalpal tibial spines and the number of teeth on the median apophysis.The South China karst, a United Nations Educational, Scientific and Cultural Organization (UNESCO) World Heritage Site since 2007, is noted for its karst features and landscapes as well as rich biodiversity. Numerous subterraean species have been reported in this region, especially invertebrate fauna . The spi species , and witLeptonetela and investigate the diversity of the genus. The standard molecular barcode, cytochrome c oxidase subunit Ⅰ (COI) was used. A species discovery method, automatic barcode gap discovery (ABGD)  to test taxonomic value of morphological characters used in traditional methods of classification.In this study, we test the usefulness of DNA barcoding for species identification in the subterranean genus y (ABGD) , and a sy (ABGD)  were botLeptonetela individuals from 122 populations (caves) (Supplementary Table S1) in Eurasia  barcode region using the primer pairs LCOI490/HCO2198 , 2.0 μL of dNTP Mix (2.5 mmol/L), 1 μL of each forward and reverse 10 μmol/L primer, 1 μL of DNA template, and 0.25 μL Taq DNA polymerase . Double-stranded PCR products were visualized by agarose gel electrophoresis (1% agarose). PCR products were purified and sequenced by Sunny Biotechnology Co., Ltd  using the ABI 3730XL DNA analyser. Sequences were aligned using ClustalW in Mega 6.0  following the manufacturer's protocol. We amplified the cytochrome c oxidase subunit Ⅰ  in RAXML v. 7.0.3 with the GTRCAT model . One hunCOI barcode dataset (see Supplementary Table S1) using two species delineation methods. DNA barcoding gap analyses require an a priori species designation. Therefore, we divided the 624 Leptonetela individuals of 122 populations (caves) into 90 putative species based on morphological characters and geographic information. In our DNA barcoding gap analysis, we examined the overlap between the mean intraspecific and interspecific Kimura 2-parameter (K2P)   and uncodistance  for eachdistance .The automatic barcode gap discovery procedure (ABGD) , which does not require assigning samples to putative species, calculates all pairwise distances in the dataset, evaluates intraspecific divergences, and then sorts the samples into candidate species using the calculated distances. We performed ABGD analyses online (http://wwwabi.snv.jussieu.fr/public/abgd/), using three different distance metrics: Jukes-Cantor (JC69) , Kimura 2-parameter (K2P) , and simple distance  . We analyzed the data using two different values for the parameters Pmin (0.0001 and 0.001), Pmax (0.1 and 0.2), and relative gap width (X=1 or 1.5), with all other parameters at default values. The terminology and the measurements in this paper generally follow This article conforms to the requirements of the amended International Code of Zoological Nomenclature. All nomenclatural acts contained within this published work have been registered in ZooBank. The ZooBank LSIDs (Life Science Identifiers) can be resolved and the associated information viewed by appending the LSID to the prefix "http://zoobank.org/". The LSID for this publication is: urn:lsid:zoobank.org:pub:7ECB1BDC-8893-4D0F-8BEA-17ECE327FC47 L. kanellisi and L. robustispina, four or more DNA barcodes were generated. All nucleotides were translated into functional protein sequences in the correct reading frame, with no stop codons or indels observed. Similar to other arthropod studies, our data indicated a high AT-content for this mitochondrial gene fragment: the mean sequence compositions were A=20.5%, C=12.6%, G=24.4%, T=41.4%.In total, 624 DNA barcodes were analyzed. A full list of the analyzed specimens can be found in Supplementary Table S1. Fragment lengths of the analyzed DNA barcodes ranged from 107 (0.005%) to 617 bp (89%). For all populations, except Leptonetela is monophyletic, with the node highly supported  whereas interspecific distances were between 3.1/3% and 31.9/25% (K2P/uncorrected p-distance). Maximum intraspecific distances > 3% were found for two species, including L. reticulopecta (4.3/4.0%), and L. pentakis Lin & Li, 2010 (5.3/5%). The lowest interspecific distance were revealed for the two species pairs L.changtu Wang & Li sp. nov. with L. chuan Wang & Li sp. nov. and L. kangsa Wang & Li sp. nov. with L. shibingensis Guo, Yu & Chen, 2016 with a value of 3.1/3%. Minimum interspecific pairwise distances < 5%, and > 3% were found for two species pairs: L. shibingensis with L. shanji Wang & Li sp. nov. and L. dao Wang & Li sp. nov. with L. xiaoyan Wang & Li sp. nov. The mean interspecific distance between the 90 tentative species was 17.9/15.6% (K2P/uncorrected p-distance), and the mean intraspecific distance within each species was 0.2% (both K2P and uncorrected p-distance) in Leptonetela. A histogram of the gap and overlap between intra-and interspecies genetic distances are show in DNA barcoding gap analysis: Based on our COI dataset, using the originally specified parameter combinations and partitions resulted mostly in 90 distinct species that correspond to the 90 species observed in the previous taxonomic hypotheses based on morphology. The result was the same regardless of the model of evolution employed (Jukes-Cantor (JC), K2P, Simple Distance). The settings P min/P max=0.0001/0.2 yielded the most significant P values. However, at lower values of prior intraspecific distance (P), recursive partitioning of ABGD recognized more species , our data represent the first published DNA barcodes.DNA barcoding is widely recognized as a useful tool for species identification across the animal kingdom . Our resCOI showed high genetic structure between populations within species .Classically, geographic isolation is considered a primary feature of troglobitic taxa . Our DNACOI sequence divergence. The interspecific genetic divergences between L. chuan Wang & Li sp. nov. and L. changtu Wang & Li sp. nov., L. kangsa Wang & Li sp. nov. and L. shibingensis, as well as between L. shibingensis and L. shanji Wang & Li sp. nov. was 3.1/3.0% based on K2P/uncorrected and p-distance models. Compared with other species L. chuan Wang & Li sp. nov., and L. changtu Wang & Li sp. nov., L. kangsa Wang & Li sp. nov., L. shibingensis and L. shanji Wang & Li sp. nov. are more closely distributed. In morphology, L. chuan Wang & Li sp. nov. and L. changtu Wang & Li sp. nov. can be distinguished by the shape of the median apophysis and the conductor ; L. kangsa Wang & Li sp. nov., L. shibingensis and L. shanji Wang & Li sp. nov. can be distinguished by the location and pattern of male pedipalpal tibial spines . Nevertheless, we found two species with maximum pairwise distance > 3%, including L. reticulopecta (specimens from Tianshegnqiao Cave are clearly distant from the rest) with 4.3/4.0%, L. pentakis (specimens from Liaoya cave is clearly distant from the rest) with 5.3/5.0%. Then we achieved a threshold of 3.11/3.0% (K2P/uncorrected and p-distance), excluding taxa from Tianshegnqiao Cave and Liaoya Cave. This threshold was interestingly close to the 3% commonly used in barcoding literature L. strinatii  1  Spermathecae thin and loosely twisted…………………………-  Not as above……………………………………………………………22  Male pedipalp with median apophysis………………………3-  Male pedipalp without median apophysis…………………93  Median apophysis like pine needles, sclerotized………4-  Not as above……………………………………………………………334  Median apophysis appears as 4 pine needle-like appendages…………………………………………………………………………5-  Median apophysis divided into more or less than 4 pine needle-shaped appendages…………………………………………6L. chakousp.nov.5  Tibia Ⅰ spine strong, conspicuous, with bifurcated tip……………………………………………………………………L. grandispina Lin & Li, 2010-  Tibia Ⅰ spine strong, located at the middle of tibia prolaterally……………………………………………6  Cymbium roughly double the length of bulb………………7-  Cymbium roughly the same length as bulb…………………8L. liuzhai Wang & Li sp. nov.7  Median apophysis divided into 15 pine needle-like appendages………………………L. shuilian Wang & Li sp. nov.-  Median apophysis divided into 2 pine needle-like appendages………………………L. pentakis Lin & Li, 20108  Cymbium constricted medially, median apophysis divided into 5 pine needle-like appendages…………………………………………………………………………………L. dao Wang & Li sp. nov.-  Cymbium not constricted medially, median apophysis divided into 2 pine needle-like appendages………………………………………………………………………………9  Male pedipalp with 5 tibial spines prolaterally…………10-  Male pedipalp with more than 5 tibial spines prolaterally…2910  Cymbium constricted and wrinkled medially……………11-  Cymbium not constricted or wrinkled medially…………2211  Tibial spines slender and without bifurcated tip…………12-  Tibial spines strong or with bifurcated tip…………………1612  Prolateral lobe tongue-shaped…………………………………13L. sanyan Wang & Li sp. nov.-  Prolateral lobe absent………13  Pedipalpal tibia with one spine significantly longer than other spines………………………………………………………………………14L. meitan Lin & Li, 2010-  Pedipalp tibia Ⅰ, Ⅱ spines nearly the same length…………………………………………………………………14  Conductor bamboo leaf-shaped in ventral view…………15L. liangfeng Wang & Li sp. nov.-  Conductor C-shaped in ventral view………………………………………………………………………L. suae Lin & Li, 201015  Embolus and conductor long, intersecting……………………………………………………………………………L. tongzi Lin & Li, 2010-  Embolus and conductor short, not intersecting………………………………………………………………………16  Pedipalpal tibia Ⅰ spine with bifurcated tip…………………17-  Pedipalpal tibia Ⅰ spine without bifurcated tip……………1917  Pedipalpal tibia Ⅰ spine strong, asymmetrically bifurcate…18L. danxia Lin & Li, 2010-  Pedipalpal tibia Ⅰ spine slender, symmetrically bifurcate………………………………………………………L. andreevi Deltshev, 198518  Pedipalpal tibia Ⅰ spine located proximally at tibia, thin spines Ⅱ, Ⅴ and Ⅵ arranged in a triangle, conductor bamboo leaf-shaped in ventral view………L. furcaspina Lin & Li, 2010-  Pedipalpal tibia Ⅰ spine located at distal 1/3 of tibia, conductor C shaped in ventral view………………………………………………………………………………19  Pedipalpal tibia Ⅰ spine longest…………………………………20-  Pedipalpal tibia Ⅱ spine longest………………………………21L. langdong Wang & Li sp. nov.20  Pedipalpal tibia Ⅰ spine bent distally, conductor reduced……………………………………………L. yaoiWang & Li, 2011-  Pedipalpal tibia Ⅰ spine not bent distally, conductor semicircular in ventral view……………………………L. lineataWang & Li, 201121  Eyes absent, pedipalpal tibia Ⅲ, Ⅴ and Ⅵ spines more slender than Ⅰ, Ⅱ spines……………L. caucasica Dunin, 1990-  Six eyes, pedipalpal tibial spines equally strong………………………………………………………………22  Conductor developed………………………………………………23-  Conductor reduced…………………………………………………2623  Pedipalpal tibia Ⅰ spine without bifurcated tip……………24L. anshun Lin & Li, 2010-  Pedipalpal tibia Ⅰ spine with bifurcated tip, other spines concentrated distally, tip of conductor bifurcated…………………………………………………………………24  Conductor bamboo leaf-shaped in ventral view…………25L. dashui Wang & Li sp. nov.-  Conductor C-shaped in ventral view, pedipalpal tibia Ⅰ spine longest………………………………L. qiangdao Wang & Li sp. nov.25  Pedipalpal tibia Ⅰ spine strong, prolateral bulbal lobe reduced………………………………………L. nuda -  Pedipalpal tibia Ⅰ spine slender, prolateral bulbal lobe tongue-shaped……………L. curvispinosa Lin & Li, 201026  Cymbium with a distal and proximal spine prolaterally, pedipalpal tibial spines equidistant……………………………………………………………………………-  Not as above……………………………………………………………27L. wangjia Wang & Li sp. nov.27  Pedipalpal tibia Ⅰ spine slender, asymmetrically bifurcated…………………………………………-  Not as above……………………………………………………………28L. maxillacostata Lin & Li, 201028  Pedipalpal tibia Ⅰ, Ⅱ, and Ⅲ spines concentrated in the mid of tibia, 2 additional spines located distally, prolateral bulbal lobe reduced……………………L. chenjia Wang & Li sp. nov.-  Pedipalpal tibia Ⅰ spine longest, located far from others, prolateral lobe small, tongue shaped…………………………………………………………………………29  Male pedipalp with 6 tibial spines retrolaterally…………30-  Male pedipalp with 7 tibial spines retrolaterally…………32L. gang Wang & Li sp. nov.30  Pedipalpal tibia Ⅰ, Ⅱ spines strong, equally length, Ⅱ spine asymmetrically bifurcated, conductor reduced………………………………………………………………-  Pedipalpal tibial spines slender, not bifurcated, conductor developed………………………………………………………………31L. gigachela 31  Pedipalpal tibia with 2 large spines prolaterally, cymbium not constricted medially, earlobe-shaped process absent, and cymbium long, twice the length of bulb…………………………………………………………………L. wenzhu Wang & Li sp. nov.-  Pedipalpal tibia without prolateral spines, cymbium constricted medially, retrolaterally attaching an earlobe-shaped process, cymbium less than twice the length of bulb…………………………………………………………………L. rudong Wang & Li sp. nov.32  Cymbium with 1 horn-shaped spine on the earlobe-shaped process, conductor thin, triangular in ventral view………………………………………………………L. la Wang & Li sp. nov.-  Earlobe-shaped process of cymbium without spine, conductor broad, C shaped in ventral view…33  Median apophysis like a pointed process or lamelliform..34-  Median apophysis finger-shaped or harrow-like………5034  Cymbium not constricted medially, earlobe-shaped process reduced……………………………………………………………………35-  Cymbium constricted medially, earlobe-shaped process developed…………………………………………………………………3835  Male pedipalpal tibia with 6 spines retrolaterally……………36-  Male pedipalpal tibia only with 5 spines retrolaterally……37L. bama Lin & Li, 201036  Pedipalpal tibia with 4 long spines prolaterally, the retrolateral Ⅰ spine longest, Ⅱ Ⅲ spines short and strong, median apophysis pointed, conductor bamboo leaf-shaped………………………………………………………………………………L. yangi Lin & Li, 2010-  Pedipalpal tibia with 3 long spines prolaterally, the retrolateral Ⅰ spine longest and strongest, median apophysis "M"-shaped, conductor reduced………………………L. liping Lin & Li, 201037  Pedipalpal tibia with 1 long spine prolaterally, the retrolateral Ⅰ spine longest and strongest, median apophysis pointed, conductor bamboo leaf-shaped………L. mayang Wang & Li sp. nov.-  Pedipalpal tibia with 3 long spines prolaterally, the retrolateral spines Ⅰ slender, and longest, median apophysis obtuse triangle shaped, conductor narrow, triangular……………………………………………………………38  Cymbium with 1 strong spine on the earlobe-shaped process…………………………………………………………………………………39-  No spine on the earlobe-shaped process…………………4339  Male pedipalp tibia with 5 spines retrolaterally…………40-  Male pedipalp tibia with more than 5 spines retrolaterally…41L. jiahe Wang & Li sp. nov.40  Cymbium with 1 curved spine retrolaterally, median apophysis pointed, with 3 sclerotized apices distally, conductor C shaped…………………………………L. panbao Wang & Li sp. nov.-  Cymbium without curved spine retrolaterally, median apophysis punctate in ventral view, conductor vestigial………………………………………………L. jiulong Lin & Li, 201041  Pedipalpal tibia Ⅰ spine strong, Ⅱ spine asymmetrically bifurcated, median apophysis lamelliform, conductor triangular…………………………………………………-  Pedipalpal tibia Ⅰ spine slender, not bifurcated…………42L. parlongaWang & Li, 201142  Pedipalpal tibia with 3 long spines prolaterally, 6 spines retrolaterally, median apophysis semicircular………………………………………………………………L. mitaWang & Li, 2011-  Pedipalpal tibia with 5 long spines prolaterally, 7 spines retrolaterally, median apophysis mita-shaped, embolus with 1 tooth distally……………………………43  Pedipalpal tibia with 5 spines retrolaterally………………44-  Pedipalpal tibia with more than 5 spines retrolaterally……4944  Pedipalpal tibia with 3 long spines prolaterally………………45-  Pedipalpal tibia with 1 or 2 long spines prolaterally………4745  Conductor C shaped in ventral view……………………………46L. xianren Wang & Li sp. nov.-  Conductor bamboo leaf-shaped in ventral view, retrolateral spines Ⅰ longest, median apophysis triangular………………………………………………………………L. rudiculaWang & Li, 201146  Pedipalpal tibia Ⅰ spine longest, the rest concentrated at distal end of tibia, median apophysis spatula-shaped in ventral view…………………………L. longli Wang & Li sp. nov.-  Pedipalpal tibia Ⅰ spine longest, Ⅰ, Ⅱ, and Ⅲ spines equally strong, median apophysis single quote shaped, " ′ " in ventral view…………………………L. pungitiaWang & Li, 201147  Pedipalpal tibia with 1 long spine prolaterally, median apophysis tongue-shaped, conductor triangular……………………………………………………………-  Pedipalpal tibia with 2 long spines prolaterally………………48L. chiosensisWang & Li, 201148  Pedipalpal tibia Ⅰ spine strongest, Ⅲ-Ⅴ spines in a triangular arrangement, median apophysis punctate, conductor triangular…………………………L. feilong Wang & Li sp. nov.-  Pedipalpal tibia Ⅰ spine longest, spine Ⅰ-Ⅲ equally strong, Ⅳ-Ⅴ situated distally median apophysis "m"-shaped, conductor triangular…………L. tiankeng Wang & Li sp. nov.49  Pedipalpal tibia with 6 spines retrolaterally, tibia Ⅰ spine close to others, median apophysis flake-like, sclerotized distally, conductor broad, undulate distally………………………………………………………………L. lophacantha -  Pedipalpal tibia with 7 spines retrolaterally, tibia Ⅰ spine distant from others, median apophysis small worm-shaped, conductor thin, triangular…………………………………………………………………50  Median apophysis index finger like…………………………51-  Median apophysis harrow-like…………………………………72L. xinhua Wang & Li sp. nov.51  Embolus bifurcated……………-  Embolus not bifurcated……………………………………………5252  Base of median apophysis swollen…………………………53-  Base of median apophysis not swollen……………………5653  Male pedipalpal tibia with 5 spines retrolaterally………54L. quinquespinata -  Male pedipalpal tibia with 6 slender spines retrolaterally, spines Ⅰ longest, conductor smooth, semicircular……………………………………………L. jinsha Lin & Li, 201054  Pedipalpal tibia Ⅰ spine much stronger than Ⅱ, asymmetrically bifurcated……………………………………-  Pedipalpal tibia Ⅰ spine similarly strong as Ⅱ, not bifurcated…………………………………………………………………………………55L. gubin Wang & Li sp. nov.55. Cymbium constricted medially, earlobe-shaped process with 2 long, curved spines retrolaterally, base of median apophysis distinctly swollen, conductor smooth, broad, semicircular…………………………L. lujia Wang & Li sp. nov.-  Cymbium not constricted medially, earlobe-shaped process small, base of median apophysis slightly swollen, conductor rugose, thin, triangular……………L. wuming Wang & Li sp. nov.56  Median apophysis bifurcated distally…………………………………………………………………………-  Median apophysis not bifurcated distally…………………5757  Pedipalpal tibia Ⅰ spine located at the base of tibia……58-  Pedipalpal tibia Ⅰ spine located medially……………………59L. shibingensis Guo, Yu & Chen, 201658  Pedipalpal tibia Ⅰ spine asymmetrically bifurcated, tibia with 4 long spines prolaterally………………………………………………………………………………L. kangsa Wang & Li sp. nov.-  Pedipalpal tibia Ⅰ spine not bifurcated…………………………………………………………………………59  Male pedipalp tibia with 6 spines retrolaterally…………60-  Male pedipalp tibia with 5 spines retrolaterally…………62L. xiaoyan Wang & Li sp. nov.60  Pedipalpal tibia with 4 spines prolaterally, cymbium with 1 curved spine at the base of retrolateral surface, median apophysis weakly sclerotized…………………………………………………………………………………-  Not as above……………………………………………………………61L. oktocantha Lin & Li, 201061  Male pedipalp tibia with 2 spines prolaterally, conductor short, broad and rugose………L. hexacantha Lin & Li, 2010-  Male Pedipalp tibia without spine prolaterally, conductor smooth, semicircular……………62  Median apophysis curved distally……………………………63-  Median apophysis not curved distally………………………65L. mengzongensisWang & Li, 201163  Cymbium with 1 horn-shaped spine on the earlobe-shaped process retrolaterally, tibia spines gradually shorted, conductor smooth, C shaped……………………………………………………………………-  Cymbium without spine on the earlobe-shaped process retrolaterally………………………………………………………………64L. hamata Lin & Li, 201064  Male pedipalp tibia with 2 long setae prolaterally, tibia Ⅰ Ⅱ and Ⅲ spines equally in length, conductor broad, semicircular…………………………………………………L. tetracantha Lin & Li, 2010-  Male pedipalp tibia with 4 long spines prolaterally, tibia Ⅰ Ⅱ spines equally in length, conductor long, curved distally……………………………………………65  Pedipalpal tibia Ⅰ spur strong……………………………………66-  Pedipalpal tibia Ⅰ spine slender…………………………………7066  Pedipalpal tibia Ⅰ spine asymmetrically bifurcated………67L. reticulopecta Lin & Li, 2010-  Pedipalpal tibia Ⅰ spine not bifurcated, conductor broad, C shaped, median apophysis distinctly sclerotized…………………………………………………………67  Median apophysis tapering……………………………………68-  Median apophysis blunt…………………………………………69L. shanji Wang & Li sp. nov.68  Pedipalpal tibia Ⅰ spine located at the middle of tibia………………………………………………………L. digitata Lin & Li, 2010-  Pedipalpal tibia Ⅰ spine located at the basal of tibia………………………………………………………………L. tianxinensis 69  Pedipalpal tibia Ⅱ-Ⅴ spines slender flexible, Ⅰ and Ⅱ spines equally length, conductor shorter than median apophysis………………………………………L. nanmu Wang & Li sp. nov.-  Pedipalpal tibia Ⅱ spine slender, Ⅲ spine strong, conductor longer than median apophysis……………………………………………………………………………………L. huoyan Wang & Li sp. nov.70  Pedipalpal tibia Ⅰ spine located at the base of tibia, other spines concentrated distally on tibia, conductor smooth, semicircular………………………-  Not as above……………………………………………………………71L. geminispina Lin & Li, 201071  Pedipalpal tibia Ⅰ Ⅱ spines adjacent, the rest short, concentrated distally, outermost plumose, tibia with 2 spines prolaterally, conductor bifurcate………………………………………………………………………L. tianxingensisWang & Li, 2011-  Pedipalpal tibia Ⅰ-Ⅳ spines spaced at regular intervals, Ⅳ and Ⅴ adjacent, tibia Ⅰ-Ⅲ equal in length, conductor short, C shaped…………………………72  Median apophysis harrow-like, horrow pin reduced to sclerotized spots………………………………………………………73-  Median apophysis harrow-like, horrow pin not reduced…75L. liuguan Wang & Li sp. nov.73  Pedipalpal tibial spines slender, equally strong, median apophysis long, half the length of bulb………………………………………………………………………-  Pedipalpal tibial spines not equally strong, median apophysis short, 1/5 the length of bulb…………………………74L. penevi Wang & Li, 201674  Pedipalpal tibia Ⅰ, Ⅱ spines equally strong, stronger than other spines, Ⅲ-Ⅴ in triangular arrangement, cymbium constricted medially, with one curved spine at the base of constriction retrolaterally…………L. changtu Wang & Li sp. nov.-  Pedipalpal tibia Ⅰ Ⅱ Ⅲ spines equally strong, stronger than other spines, Ⅲ-Ⅴ not triangular arrangement, cymbium not constricted medially…………75  Median apophysisi harrow-like, the horrow pin not constant in size………………………………………………………………………76-  Median apophysisi harrow-like, the horrow pin constant in size…………………………………………………………………………8076  Pedipalpal tibia Ⅰ spine not bifurcated…………………………77L. lianhua Wang & Li sp. nov.-  Pedipalpal tibia Ⅰ spine strong, asymmetrically bifurcated, other 4 spines slender, median apophysis with 5 small teeth and 1 large, horn-shaped tooth…………………………………………………………………………77  Pedipalpal tibia Ⅰ spine longest…………………………………78-  Pedipalpal tibia Ⅱ spine longest………………………………79L. megaloda 78  Median apophysis palmate, with six teeth distally…………………………………………L. niubizi Wang & Li sp. nov.-  Median apophysis antler-like, with 4 small teeth and 1 large tooth, which bears 2 small teeth…………………………………………………………………………………L. hangzhouensis 79  Two large teeth on the periphery of median apophysis, 2 small teeth in the middle……………………………………………………………………L. microdonta -  Two large teeth on the periphery of median apophysis, 5 small teeth in the middle.. 80  Median apophysis short and broad…………………………81-  Median apophysis long and thin……………………………8781  Pedipalpal tibia Ⅰ spine strongest……………………………82-  Pedipalpal tibia Ⅰ spine not strongest………………………83L. identica 82  Pedipalpal tibia Ⅰ, Ⅱ spines equally strong, stronger than other 3 spines, median apophysis with 6 small teeth apically………………………………L. meiwang sp.nov.-  Pedipalpal tibia Ⅱ, Ⅲ spines equally strong, spine Ⅱ longest, median apophysis with 5 sharp teeth apically……………………………………………………………………………83  Pedipalpal tibia Ⅰ spine bifurcated……………………………84-  Pedipalpal tibia Ⅰ spine not bifurcated………………………8984  Distal edge of median apophysis with 6 teeth……………85-  Distal edge of median apophysis with 5 or 10 teeth…86L. zakou Wang & Li sp. nov.85  Teeth of median apophysis needle-shaped, earlobe-shaped process of cymbium absent; in the female, anterior margin of atrium with one pointed process medially…………………………………………………………………L. sexdentataWang & Li, 2011-  Teeth of median apophysis normal, cymbium with earlobe-shaped process, female anterior margin of atrium without pointed process………………L. longyu Wang & Li sp. nov.86  Distal edge of median apophysis with 5 teeth, conductor C shaped, tip of conductor undulate in the female, anterior margin of atrium with one pointed process medially…………………………………L. shicheng Wang & Li sp. nov.-  Distal edge of median apophysis with 10 teeth, conductor C shaped, distal edge of conductor smooth; in the female, anterior margin of atrium without pointed process…………………………………………………87  Pedipalpal tibia Ⅱ spine tapering………………………………88L. flabellarisWang & Li, 2011-  Pedipalpal tibia Ⅰ spine blunt…L. palmata Lin & Li, 201088  Distal edge of median apophysis with 5 teeth, conductor short, C shaped………………………L. kanellisi -  Distal edge of median apophysis with 7 teeth, conductor long, triangular shaped………………………………………………………………………………89  Pedipalpal tibia with clusters of short spines dorsally……90-  Pedipalpal tibia without clusters of short spines dorsally………………………………………………………………………………………91L. encun Wang & Li sp. nov.90  Distal edge of median apophysis linear, with 8 teeth……………………………………………………L. robustispina -  Distal edge of median apophysis semicircular, with 12 teeth………………………91  Base of pedipalpal tibia swollen………………………………92-  Base of pedipalpal tibia not swollen…………………………9692  Pedipalpal tibia Ⅰ spine bifurcate………………………………93L. notabilis -  Pedipalpal tibia Ⅰ spine trifurcate…………………………………………………………………………………93  Conductor triangular, longer than median apophysis, median apophysis with 7 teeth………………………………………………94-  Conductor C shaped, shorter than median apophysis, median apophysis with 6 teeth……………………………………95L. shuang Wang & Li sp. nov.94  Spermathecae not twisted distally………………………………………………………………………………L. sanchahe Wang & Li nom. nov.-  Spermathecae twisted distally……………………………………………………………………………L. sexdigiti 95  Spermathecae weakly twisted……………………………………………………………………………L. lihu Wang & Li sp. nov.-  Spermathecae strongly twisted………………………………………………………………………………………96  Pedipalpal tibia Ⅰ spine strongset……………………………97-  Pedipalpal tibia Ⅰ, Ⅱ, Ⅲ spines equally strong, stronger than other spines……………………………………………………………10397  Pedipalpal tibia Ⅱ spine bifurcate………………………………98-  Pedipalpal tibia Ⅰ spine not bifurcate………………………10198  Pedipalpal tibia Ⅱ-Ⅴ spine slender, curved, equally strong…………………………………………………………………………………99-  Pedipalpal tibia Ⅱ-Ⅴ spine not equally strong…………100L. arvanitidisi Wang & Li, 201699  Distal edge of median apophysis with 6 teeth……………………………………………………L. erlong Wang & Li sp. nov.-  Distal edge of median apophysis with 5 teeth………………………………………………………L. tawo Wang & Li sp. nov.100  Distal edge of median apophysis with 4 teeth, tibia Ⅱ, Ⅲ spines equally strong, stronger than other 2 spines………………………………………………L. paragamiani Wang & Li, 2016-  Distal edge of median apophysis with 3 teeth, tibia Ⅲ-Ⅴ spines equally strong, slender than spine Ⅱ……………………………………………………………101  Pedipalpal tibia Ⅱ-Ⅴ spines equally strong………………102L. deltshevi -  Pedipalpal tibia Ⅲ-Ⅴ spines equally strong, slender than spine Ⅱ, distal edge of median apophysis with 4 teeth……………………………………………………L. gittenbergeriWang & Li, 2011102  Distal edge of median apophysis with 5 teeth, conductor C shaped…………………………L. zhaiWang & Li, 2011-  Distal edge of median apophysis with 6 teeth, conductor semicircular………………………………L. thracia Gasparo, 2005103  Pedipalpal tibia Ⅰ Ⅱ spines equally strong……………………………………………………………………………-  Pedipalpal tibia Ⅱ and Ⅲ spines equally strong………104L. dabian Wang & Li sp. nov.104  Distal edge of median apophysis with 3 teeth, tibia with 3 large spines prolaterally………L. chuan Wang & Li sp. nov.-  Distal edge of median apophysis with 6 teeth, tibia with 6 long setae prolaterally…………Type species:Leptonetela kanellisi  from Greece.Diagnosis. The genus Leptonetela can be distinguished from other leptonetid genera by the following combination of male pedipalpal characters: femur lacking spines and tibia with a longitudinal row of spines on the retrolateral surface.Redescription. Carapace yellowish or white. Sternum shield-shaped. Opisthosoma gray, ovoid, covered with short hairs. Male pedipalpal patella with one short spine dorso-distally; tibia with trichobothria dorsally; cymbium with strong, thorny spine distally; bulb yellowish, ovoid, with two appendages inserted ventrally, median apophysis chitinous, conductor membranous, median apophysis and conductor absent in some species, embolus transparent, membranous. Female genital area covered with short hairs. Vulva with a pair of spermathecae and sperm ducts, spermathecae twisted and weakly sclerotized.Distribution. Greece, Turkey, Georgia, Azerbaijan, Vietnam and China.Type material. Holotype: male (IZCAS), Chakou Cave, N27.93°, E106.14°, Shalang, Shibao Town, Gulin County, Luzhou City, Sichuan Province, China, 20 April 2014, Y. Li, H. Zhao & Y. Lin leg. Paratypes: 1 male and 3 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. dao Wang & Li sp. nov., L. grandispina Lin & Li, 2010, L. liuzhai Wang & Li sp. nov., L. pentakis Lin & Li, 2010, and L. shuilian Wang & Li sp. nov., but can be distinguished by the male pedipalpal tibia with 5 spines retrolaterally, the basal spine strong, conspicuous and with a bifurcate tip ; the median apophysis divided into 4 pine needle like structures ; from L. dao Wang & Li sp. nov., L. grandispina, L. pentakis by the conductor reduced in this new species (L. liuzhai Wang & Li sp. nov. by the cymbium 1.3 times longer than bulb (L. liuzhai Wang & Li sp. nov. and L. shuilian Wang & Li sp. nov.).cate tip  . Total length 2.25 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.27 (gth 2.27 -B. Carapgth 2.27 : spermatDistribution. China (Sichuan).Type material. Holotype: male (IZCAS), Dao Cave, N27.19°, E105.06°, Shuanglong, Salaxi County, Bijie City, Guizhou Province, China, 18 November 2011, H. Chen & Z. Zha leg. Paratypes: 1 male and 20 females, same data as holotype; 5 males and 7 females, Shanlanqiao Cave, N26.28°, E106.04°, Shanlanqiao, Qianyanqiao Town, Anshun City, Guizhou Province, China, 4 November 2011, H. Chen & Z. Zha leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. chakou Wang & Li sp. nov., L. grandispina Lin & Li, 2010, L. liuzhai Wang & Li sp. nov.L. pentakis Lin & Li, 2010, and L. shuilian Wang & Li sp. nov., but can be separated from L. chakou Wang & Li sp. nov., L. grandispina, L. liuzhai Wang & Li sp. nov. and L. pentakis by median apophysis divided into 2 pine needlelike ; from L. chakou Wang & Li sp. nov., L. grandispina by the tibial spines slender (L. chakou Wang & Li sp. nov. and Ⅱ spines in L. grandispina strong); from L. chakou Wang & Li sp. nov., L. pentakis by the cymbium not constricted medially (L. liuzhai Wang & Li sp. nov. and L. shuilian Wang & Li sp. nov. by the cymbium 1.2 times longer than bulb (L. liuzhai Wang & Li sp. nov. and L. shuilian Wang & Li sp. nov.).edlelike  . Total length 2.28 . Similar to male in color and general features, but larger and with longer legs. Total length 2.76 (gth 2.76 -B. Carapgth 2.76 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), nameless Cave, N25.27°, E107.43°, Longli, Liuzhai Town, Nandan County, Hechi City, Guangxi Zhuang Autonomous Region, China, 29 January 2015, Y. Li & Z. Chen leg. Paratypes: 2 males and 6 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. chakou Wang & Li sp. nov., L. dao Wang & Li sp. nov., L. grandispina Lin & Li, 2010, L. pentakis Lin & Li, 2010, and L. shuilian Wang & Li sp. nov. but can be separated from L. chakou Wang & Li sp. nov., L. dao Wang & Li sp. nov., L. grandispina, and L. pentakis by the male pedipalpal cymbium double the length of bulb, median apophysis divided into 15 pine needle-like structures ; from L. chakou Wang & Li sp. nov., and L. grandispina by the tibial spines slender (L. chakou Wang & Li sp. nov. and Ⅱ spines in L. grandispina strong); from L. chakou Wang & Li sp. nov., and L. pentakis by the cymbium not constricted medially in this new species . Total length 2.25 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.50 (gth 2.50 -B. Carapgth 2.50 : spermatDistribution. China (Guangxi).Type material. Holotype: male (IZCAS), Shuilian Cave, N24.43°, E106.97°, Pingle, Fengshan County, Hechi City, Guangxi Zhuang Autonomous Region, China, 22 March 2015, Y. Li & Z. Chen leg. Paratypes: 6 males and 4 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. chakou Wang & Li sp. nov., L. dao Wang & Li sp. nov., L. grandispina Lin & Li, 2010, L. pentakis Lin & Li, 2010, and L. liuzhai Wang & Li sp. nov. but can be separated from L. chakou Wang & Li sp. nov., L. dao Wang & Li sp. nov., L. grandispina Lin & Li, 2010, L. pentakis Lin & Li, 2010 by the male pedipalpal cymbium double the length of bulb; from L. chakou Wang & Li sp. nov., L. grandispina, L. liuzhai Wang & Li sp. nov. and L. pentakis by the median apophysis divided into 2 pine needlelike structures in L. chakou Wang & Li sp. nov. ; from L. chakou Wang & Li sp. nov. and L. grandispina by the tibial spines slender (L. chakou Wang & Li sp. nov. and Ⅱ spines in L. grandispina strong); from L. chakou Wang & Li sp. nov. and L. pentakis by the cymbium not constricted medially in this new species.sp. nov.  . Total length 2.25 . Similar to male in color and general features, but smaller and with shorter legs. Total length 2.10 (gth 2.10 -B. Carapgth 2.10 : spermatDistribution. China (Guangxi).Type material. Holotype: male (IZCAS), Chenjia Cave, N28.38°, E108.67°, Tianba, Songtao County, Tongren City, Guizhou Prvince, China, 9 March 2013, H. Zhao & J. Liu leg. Paratypes: 1 male and 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. anshun Lin & Li, 2010, L. suae Lin & Li, 2010, L. tongzi Lin & Li, 2010, L. meitan Lin & Li, 2010, L. liangfeng Wang & Li sp. nov., and L. sanyan Wang & Li sp. nov., but can be distinguished by the male pedipalal tibia Ⅰ spine far apart from the other 4 spines ; is also similar to L. huoyan Wang & Li sp. nov., but can be distinguished by the absent median apophysis, reduced conductor (L. huoyan Wang & Li sp. nov.).4 spines , conduct4 spines  . Total length 2.50 . Similar to male in color and general features, but smaller and with shorter legs. Total length 2.25 (gth 2.25 -B. Carapgth 2.25 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Liangfeng Cave, N28.32°, E107.84°, Tian, Fengle Town, Wuchuan County, Zunyi City, Guizhou Province, China, 7 August 2012, H. Zhao leg. Paratypes : 1 male and 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. anshun Lin & Li, 2010, L. suae Lin & Li, 2010, L. tongzi Lin & Li, 2010, L. meitan Lin & Li, 2010, L. chenjia Wang & Li sp. nov., and L. sanyan Wang & Li sp. nov., but can be distinguished by the male pedipalpal bulb conductor C shaped ; from L. anshun by the tibia Ⅰ spine slender not bifurcated (L. anshun).C shaped  . Total length 2.28 . Similar to male in color and general features, but smaller and with shorter legs. Total length 2.14 (gth 2.14 -B. Carapgth 2.14 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Sanyan Cave, N29.15°, E107.60°, Heyi, Yangxi Town, Daozhen County, Guizhou Province, China, 30 May 2011, Z. Zha leg. Paratypes: 1 male and 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. anshun Lin & Li, 2010, L. suae Lin & Li, 2010, L. tongzi Lin & Li, 2010, L. meitan Lin & Li, 2010, L. chenjia Wang & Li sp. nov., and L. liangfeng Wang & Li sp. nov., but can be separated from all above except L. tongzi by in the male conductor C shaped in this new species ; from L. tongzi by in the female atrium triangular, anterior margin of the atrium undulate (L. tongzi). species  . Total length 1.78 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.03 (gth 2.03 -B. Carapgth 2.03 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Wangjia Cave, N26.98°, E107.94°, Gaoqi, Nongchang Town, Huangpin County, Guizhou Province, China, 4 March 2012, H. Zhao & J. Liu leg. Paratype: 1 female, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. danxia Lin & Li, 2010, and L. yaoiL. danxia, bamboo leaf-shaped in L. yaoi), from L. yaoi by the tibia Ⅰ spine slender, asymmetrically bifurcated (L. yaoi); from L. danxia by the unwrinkled cymbium (L. danxia).furcated  . Total length 2.13 .Type material. Holotype: male (IZCAS), Qiangdao Cave, N25.83º, E109.04º, Guandong Town, Congjiang County, Qiandongnan Prefecture, Guizhou, China, 16 March 2013, H. Zhao & J. Liu leg. Paratypes: 3 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. furcaspina Lin & Li, 2010, L. langdong Wang & Li sp. nov. and L. dashui Wang & Li sp. nov., but can be distinguished by the male pedipalpal tibia Ⅰ spine strong .e strong , conducte strong  . Total length 1.80 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.25 (gth 2.25 -B. Carapgth 2.25 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Menglonggong Cave, N27.07°, E107.76°, Langdong Village, Huangping County, Qiandongnan Prefecture, Guizhou Province, China, 3 March 2013, H. Zhao & J. Liu leg. Paratypes: 1 male and 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. furcaspina Lin & Li, 2010, L. qiangdao Wang & Li sp. nov., and L. dashui Wang & Li sp. nov., but can be distinguished by the male pedipalpal tibia Ⅰ spine strong, tip curved .p curved . conductp curved  : total length 2.25 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.25 (gth 2.25 -B. Carapgth 2.25 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Dashui Cave, N26.61°, E106.61°, Shijicheng, Jinyang New Urban Area, Guiyang City, Guizhou Province, China, 18 June 2011, Z. Zha leg. Paratypes: 1 male and 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. furcaspina Lin & Li, 2010, L. qiangdao Wang & Li sp. nov., and L. langdong Wang & Li sp. nov., but can be distinguished by the slender male pedipalpal tibia Ⅰ spine, Ⅱ Ⅲ spines curved basally . basally , conduct basally , . Total length 1.88 . Similar to male in color and general features, but larger and with shorter legs. Total length 1.93 (gth 1.93 -B. Carapgth 1.93 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Gang Cave, N26.87°, E108.91°, Tunhou, Nanming Town, Jianhe County, Kaili City, Guizhou Province, China, 15 December 2011, Z. Zha leg. Paratypes: 15 males and 6 females, same data as holotype; 4 males and 5 females, Long Cave, N26.85°, E108.79°, Longtang, Liangshang Town, Sansui County, Kaili City, Guizhou Province, China, 18 December 2011, Z. Zha leg; 5 males and 5 females, Shenxian Cave, N26.87°, E108.89°, Shixing, Xiaolan Country, Nanming Town, Jianhe County, Kaili City, Guizhou Province, China, 16 December 2011, Z. Zha leg; 5 females, Niu Cave, N26.86°, E108.93°, Cenge, Nanming Town, Jianhe County, Kaili City, Guizhou Province, China, 14 December 2011, Z. Zha leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. jiulong Lin & Li, 2010, but can be distinguished by the male pedipalpal tibia with 6 spines retrolaterally, tibia Ⅱ spine thickest, Ⅰ, Ⅱ spines equally length, tibia Ⅱ spine asymmetrically bifurcated (L. jiulong).furcated , median furcated  . Total length 2.63 . Similar to male in color and general features, but smaller and with shorter legs. Total length 2.38 (gth 2.38 -B. Carapgth 2.38 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), La Cave in Xiaoyakou, N25.80°, E104.95°, Puan County, Qianxinan Prefecture, Guizhou Province, China, 14 July 2012, H. Zhao leg. Paratypes: 3 males and 5 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. rudong Wang & Li sp. nov. and L. wenzhu Wang & Li sp. nov. but can be distinguished from L. wenzhu Wang & Li sp. nov. by the male pedipalpal tibia with 7 spines retrolaterally ; from L. rudong Wang & Li sp. nov. by the tibia with 4 long setae prolaterally (L. rudong Wang & Li sp. nov.); from L. rudong Wang & Li sp. nov., and L. wenzhu Wang & Li sp. nov. by the conductor broad, C shaped .aterally  . Total length 2.97 . Similar to male in color and general features, but smaller and with shorter legs. Total length 2.81 (gth 2.81 -B. Carapgth 2.81 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Rudong Cave, N25.57°, E110.62°, Longpan Mountain, Dongtian, Xing'an County, Guilin City, Guangxi Zhuang Autonomous Region, China, 11 July 2009, C. Wang & Z. Yao leg. Paratypes: 1 male and 3 females, same data as holotype; 3 females, Gouya Cave, N25.46°, E110.11°, Hufeng, Guanyang County, Guilin City, Guangxi Zhuang Autonomous Region, China, 30 August 2009, C. Wang & Z. Yao leg; 2 females, Jiulong Cave, N25.46°, E110.09°, Shifeng, Guanyang County, Guilin City, Guangxi Zhuang Autonomous Region, China, 30 August 2009, C. Wang & Z. Yao leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. la Wang & Li sp. nov., and L. wenzhu Wang & Li sp. nov. but can be distinguished by the male pedipalpal tibia with 2 long setae, 2 spines prolaterally, 1 long seta, and 6 spines retrolaterally, cymbium with 1 horn-shaped spine on the earlobe-shaped process . process -D, condu process  . Total length 2.12 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.15 , Wenzhu Cave, N25.44°, E105.13°, Longchang Town, Xingren City, Guizhou Province, China, 16 July 2012, H. Zhao leg. Paratypes: 1 male and 2 females, same data as holotype; 4 females, Xiaoya Cave, N25.44°, E105.13°, Yaqiao Town, Xingren City, Guizhou Province, China, 16 July 2012, H. Zhao leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. rudong Wang & Li sp. nov., and L. la Wang & Li sp. nov. but can be distinguished by the male pedipalpal tibia with 6 spines retrolaterally .aterally , conductaterally  . Total length 2.63 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.88 (gth 2.88 -B. Carapgth 2.88 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Underground River, N25.27°, E107.44°, Longli, Liuzhai Town, Nandan County, Hechi City, Guangxi Zhuang Autonomous Region, China, 29 January 2015, Y. Li & Z. Chen leg. Paratypes: 3 males and 4 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. chiosensisL. panbao Wang & Li sp. nov., but can be distinguished by the male pedipalpal tibia Ⅰ, Ⅱ and Ⅲ spines equally strong .y strong , conducty strong  . Total length 1.88 . Similar to male in color and general features, but larger and with shorter legs. Total length 1.95 (gth 1.95 -B. Carapgth 1.95 : spermatDistribution. China (Guangxi).Type material. Holotype: male (IZCAS), Panbao Cave, N28.38°, E108.67°, Panbao, Shichang Town, Songtao County, Tongren City, Guizhou Province, China, 8 March 2013, H. Zhao & J. Liu leg. Paratypes: 2 male and 4 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. chiosensisL. Longli Wang & Li sp. nov., but can be distinguished by the male pedipalpal tibial spines slender, equally strong, cymbium with 1 strong spine on the earlobe-shaped process . process , conduct process  . Total length 2.38 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.50 (gth 2.50 -B. Carapgth 2.50 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Feilong Cave, N26.44°, E107.02°, Longli Town, Qiannan Prefecture, Guizhou Province, China, 27 July 2012, H. Zhao leg. Paratypes: 9 females, same data as holotype; 1 female, Lianhua Cave, N26.43°, E106.95°, Lianhua Town, Qiannan Prefecture, Guizhou Province, China, 27 July 2012, H. Zhao leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. yangi Lin & Li, 2010 and L. jiahe Wang & Li sp. nov., but can be distinguished from L. yangi by the male pedipalpal cymbium constricted medially, attaching an earlobe-shaped process (L. yangi), from L. jiahe Wang & Li sp. nov. by the median apophysis "m"-shaped, conductor triangular . process , conduct process  . Total length 2.31 . Similar to male in color and general features, but smaller and with shorter legs. Total length 2.13 (gth 2.13 -B. Carapgth 2.13 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Jiahe Cave, N25.25°, E110.20°, Lingui Town, Lingui County, Guilin City, Guangxi Zhuang Autonomous Region, China, 20 December 2013, H. Zhao leg. Paratypes: 3 males and 5 females, same data as holotype; 6 males and 5 females, Flytiger Cave, N25.25°, E110.20°, Lingui Town, Lingui County, Guilin City, Guangxi Zhuang Autonomous Region, China, 20 December 2013, H. Zhao leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. yangi Lin & Li, 2010, and L. feilong Wang & Li sp. nov., but can be distinguished by the male pedipalpal cymbium with 1 short spine on the earlobe-shaped process, and 1 curved, long spine retrolaterally .aterally , median aterally  . Total length 2.43 : tibia with 5 slender spines retrolaterally, spines Ⅰ, Ⅱ and Ⅲ equally strong, stronger than other spines, spines Ⅰ longest. Cymbium constricted medially, retrolaterally attaching to 1 curved spine and an earlobe-shaped process, with 1 short spine. Embolus triangular, and prolateral lobe absent. Median apophysis shaped like pointed process, with 3 sclerotized spots distally. Conductor C shaped . Similar to male in color and general features, but smaller and with shorter legs. Total length 2.30 (gth 2.30 -B. Carapgth 2.30 : spermatDistribution. China (Guangxi).Type material. Holotype: male (IZCAS), Xianren Cave, N29.73°, E110.31°, Yvpingxini, Zouma Town, Hefeng County, Enshi Tujia and Miao Autonomous Prefecture, Hubei Province, China, 27 January 2011, Y. Li & J. Liu leg. Paratypes: 2 males and 3 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. liping Lin & Li, 2010, and L. parlongaL. liping, 6 slender spines in L. parlonga); median apophysis triangular ; from L. parlonga by the cymbium retrolaterally with 1 horn-shaped spine on the earlobe-shaped process in L. parlonga.iangular  . Total length 2.23 . Similar to male in color and general features but larger and with shorter legs. Total length 2.38 (gth 2.38 -B. Carapgth 2.38 : spermatDistribution. China (Hubei).Type material. Holotype: male (IZCAS), Tiankeng Cave, N26.64°, E104.80°, Hegou, Dewu Town, Zhongshan County, Liupanshui City, Guizhou Province, China, 9 November 2011, H. Chen & Z. Zha leg. Paratypes: 4 males and 5 females, same data as holotype; 2 females, Luoshui Cave, N26.64°, E104.80°, Hegou, Dewu Town, Zhongshan County, Liupanshui City, Guizhou Province, China, 9 November 2011, H. Chen & Z. Zha leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. rudiculaL. rudicula).Description. Male (holotype). Total length 2.03 . Similar to male in color and general features, but smaller and with shorter legs. Total length 1.93 (gth 1.93 -B. Carapgth 1.93 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Mayang Cave, N28.55°, E108.06°, Quankou, Dejiang County, Tongren City, Guizhou Province, China, 10 August 2012, H. Zhao leg. Paratype: 1 female, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species can be distinguished from all other species of the genus by the male pedipalpal cymbium with one curved, short spine medially in retrolateral view, median apophysis triangular, spermathecae not tightly twisted, just spiraled in the female.Description. Male (holotype). Total length 2.13 .Type material. Holotype: male (IZCAS), Gubin River, N26.50°, E107.52°, Gubin, Xingshan Town, Majiang County, Shengkaili City, Guizhou Province, China, 28 November 2011, H. Chen & Z. Zha leg. Paratypes: 22 males and 14 females, same data as holotype; 4 males and 5 females, nameless Cave, N26.50°, E107.52°, Gubin, Xingshan Town, Majiang County, Shengkaili City, Guizhou Province, China, 28 November 2011, H. Chen & Z. Zha leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. jinsha Lin & Li, 2010, L. quinquespinata , L. xinhua Wang & Li sp. nov., L. lujia Wang & Li sp. nov. and L. xinhua Wang & Li sp. nov. but can be distinguished by the male pedipalpal tibia with 4 slender spines prolaterally, 5 slender spines retrolaterally, with spines Ⅰ, Ⅱ equal length, cymbium with 2 long curved spines on earlobe-shaped process retrolaterally ; from L. jinsha, L. lujia Wang & Li sp. nov. and L. xinhua Wang & Li sp. nov. by the semicircular conductor, base of median apophysis distinctly swollen, 4 times wider than the tip .aterally  . Total length 1.88 . Similar to male in color and general features, but larger and with longer legs. Total length 2.30 (gth 2.30 -B. Carapgth 2.30 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Wuming Cave, N26.48º, E107.54º, Lujia Bridge, Gubin, Xingshan Town, Majiang County, Kaili City, Guizhou Province, China, 29 November 2011, H. Chen & Z. Zha leg. Paratypes: 1 male and 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. jinsha Lin et Li, 2010, L. quinquespinata , L. xinhua Wang & Li sp. nov. and L. gubin Wang & Li sp. nov. but can be distinguished by the male pedipalpal tibia with 4 long setae prolaterally, 5 slender spines retrolaterally, with spines Ⅰ longest, spines Ⅱ Ⅲ equal length ; from L. gubin and L. quinquespinata by the slightly swollen base of the median apophysis .l length , conductl length , . Total length 1.72 . Similar to male in color and general features, but larger and with shorter legs. Total length 1.70 (gth 1.70 -B. Carapgth 1.70 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), nameless Cave, N27.85º, E111.31º, Caojia Town, Xinhua County, Loudi City, Hunan Province, China, 24 March 2016, Y. Li & Z. Chen leg. Paratypes: 3 males and 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. jinsha Lin & Li, 2010, L. quinquespinata , L. lujia Wang & Li sp. nov., and L. gubin Wang & Li sp. nov., but can be distinguished by the bifurcated embolus, male pedipalpal tibia with 5 slender spines prolaterally, 5 slender spines retrolaterally, conductor triangular ; from L. gubin and L. quinquespinata by the base of median apophysis slightly swollen .iangular , : total length 1.78 : similar to male in color and general features, but with a larger body size and shorter legs. Total length 1.95 (gth 1.95 -B. Prosogth 1.95 : spermatDistribution. China (Hunan).Type material. Holotype: male (IZCAS), Kangsagulie Cave, N26.79º, E108.21º, Datang, Geyi Town, Taijiang County, Kaili City, Guizhou Province, China, 5 December 2011, H. Chen & Z. Zha leg. Paratypes: 7 males and 6 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. shibingensis and L. wuming Wang & Li sp. nov., but can be distinguished by the median apophysis index finger-like in prolaterally view, tip bifurcated ; from L. wuming Wang & Li sp. nov., by the tibia Ⅰ spine located at the middle of tibia, embolus with 1 basal tooth .furcated  . Total length 2.07 . Similar to male in color and general features, but smaller and with shorter legs. Total length 2.02 (gth 2.02 -B. Carapgth 2.02 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Wuming Cave, N25.43°, E105.62°, Dabei Town, Zhenfeng County, Guizhou Province, China, 18 July 2012, H. Zhao leg. Paratypes: 2 males and 5 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. kangsa Wang & Li sp. nov., and L. shibingensis but can be distinguished by the male pedipalpal bulb embolus without basal tooth (L. kangsa Wang & Li sp. nov. and L. shibingensis by the tibia spines Ⅰ located at the base of the tibia (L. kangsa Wang & Li sp. nov. and L. shibingensis).al tooth , . Total length 1.50 . Similar to male in color and general features, but larger and with shorter legs. Total length 3.02 (gth 3.02 -B. Carapgth 3.02 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Shanji Cave, N27.28°, E107.82°, Xiaguihua, Xiaosai Town, Yuqing County, Zunyi City, Guizhou Province, China, 15 August 2012, H. Zhao leg. Paratypes: 3 males and 3 females, same data as holotype; 2 females, Guanyin Cave, N27.32°, E107.71°, Hongjun, Longxi Town, Yuqing County, Zunyi City, Guizhou Province, China, 15 August 2012, H. Zhao leg; 3 females, Liangfeng Cave, N27.27°, E107.76°, Xiaosai Town, Yuqing County, Zunyi City, Guizhou Province, China, 14 August 2012, H. Zhao leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. digitata Lin & Li, 2010, L. hamata Lin & Li, 2010 and L. tetracantha Lin & Li, 2010, but can be distinguished by the male pedipalpal tibia spines Ⅰ strong, located medially ; from L. digitata by themedian apophysis not curved (median apophysis curved in L. digitata).medially  . Total length 2.08 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.40 (gth 2.40 -B. Carapgth 2.40 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Gejiaxiaoyan Cave, N27.11°, E105.24°, Shanjiao, Zhuchang Town, Bijie City, Guizhou Province, China, 27 January 2011, H. Chen & Z. Zha leg. Paratypes: 2 males and 6 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. curvispinosa Lin & Li, 2010, but can be distinguished by the male pedipalpal tibia with 4 large spines prolaterally, 6 large spines retrolaterally (L. curvispinosa).aterally , median aterally  . Total length 1.67 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.12 (gth 2.12 -B. Carapgth 2.12 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Heyuantou nameless Cave, N29.25°, E109.35°, Huoyan Street, Guitang Dam Town, Longshan County, Hubei Province, China, 15 January 2014, Y. Li & Y. Lin leg. Paratypes: 1 male and 2 females, same data as holotype; 1 male and 4 females, nameless Cave, N29.61°, E109.17° Jieping, Xianfeng County, Enshi Tujia and Miao Autonomous Prefecture, Hubei Province, China, 17 January 2014, Y. Li & Y. Lin leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. anshun Lin & Li, 2010 and L. chenjia Wang & Li sp. nov., but can be distinguished by on the male pedipalpal bulb median apophysis slightly sclerotized, index finger like, conductor broad, semicircular ; from L. anshun by the tibia spines Ⅰ slender (L. anshun).circular  . Total length 2.25 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.53 (gth 2.53 -B. Carapgth 2.53 : spermatDistribution. China (Hubei).Type material. Holotype: male (IZCAS), Liuguan Cave, N26.15°, E106.46°, Mengqiu, Baiyunshan Town, Changshun County, Guizhou Province, China, 23 December 2010, Z. Zha & Z. Chen leg. Paratypes: 2 female, same data as holotype; 1 male, Fenghuang Cave, N26.09°, E106.39°, Shenglian, Zhonghuo Town, Changshun County, Guizhou Province, China, 23 December 2010, Z. Zha & Z. Chen leg.Etymology. The specific name refers to the typelocality; noun.Diagnosis. This new species is similar to L. penevi Wang & Li, 2016 and L. changtu Wang & Li sp. nov. but can be distinguished by on the male pedipalpal bulb median apophysis long, and half the length of bulb ; male pedipalpal tibia spines slender, equally strong . of bulb  . Total length 1.88 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.08 (gth 2.08 -B. Carapgth 2.08 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Nanmu Cave, N28.10°, E110.08°, Pushi Town, Luxi County, Hunan Province, China, 5 April 2016, Y. Li & Z. Chen leg. Paratypes: 3 males and 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. tianxingensis, but can be distinguished by on the male pedipalpal bulb conductor longer than median apophysis (L. tianxingensis); male pedipalpal tibia Ⅲ spine strong (L. tianxingensis).pophysis  : total length 1.70 : similar to male in color and general features, but with a larger body size and longer legs. Total length 1.98 (gth 1.98 -B. Prosogth 1.98 : spermatDistribution. China (Hunan).Type material. Holotype: male (IZCAS), Changtu Cave, N27.14°, E105.43°, Honglin, Qianxi Town, Bijie County, Guizhou Province, China, 23 November 2011, Z. Zha & Z. Zha leg. Paratypes: 1 male and 10 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. penevi Wang & Li, 2016 and L. liuguan Wang & Li sp. nov. but can be distinguished by the male pedipalpal tibia spines Ⅰ, Ⅱ, Ⅲ equally strong, stronger than other two spines ; from L. liuguan Wang & Li sp. nov. by median apophysis short, 1/5 the length of bulb (L. liuguan Wang & Li sp. nov.); from L. penevi by the cymbium not constricted .o spines  . Total length 2.33 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.70 (gth 2.70 -B. Carapgth 2.70 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Lianhua Cave, N25.48°, E114.09°, Niedou Town, Chongyi County, Jiangxi Province, China, 24 April 2013, Y. Luo & J. Liu leg. Paratypes: 3 males and 10 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. niubizi Wang & Li sp. nov. but can be distinguished by the male pedipalpal tibia with 5 spines retrolaterally, with Ⅰ spine strongest, tip bifurcated, the other 4 spines slender, 2 of them longer than Ⅰ spine (L. niubizi Wang & Li sp. nov.). Ⅰ spine ; tip of  Ⅰ spine  . Total length 2.00 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.25 (gth 2.25 -B. Carapgth 2.25 : spermatDistribution. China (Jiangxi).Type material. Holotype: male (IZCAS), Niubizi Cave, N27.62°, E106.67°, Leshan Town, Zunyi County, Zunyi City, Guizhou Province, China, 1 August 2012, H. Zhao leg. Paratypes: 7 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. lianhua Wang & Li sp. nov. but can be distinguished by the male pedipalp tibia with 5 slender spines retrolaterally, spines Ⅰ longest, not bifurcated (L. lianhua Wang & Li sp. nov.).furcated , median furcated  . Total length 2.53 . Similar to male in color and general features, but with a larger body size and shorter legs. Total length 2.60 (gth 2.60 -B. Carapgth 2.60 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Longyu Cave, N29.40°, E110.09°, Cili County, Hunan Province, China, 5 June 2011, Z. Zha leg. Paratypes: 4 males and 5 females, same data as holotype, 5 males and 6 females, Niuerduo Cave, N29.404°, E110.73°, Cili County, Hunan Province, China, 9 April 2016, Y. Li & Z. Chen leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. sexdentataL. shicheng Wang & Li sp. nov., L. zakou Wang & Li sp. nov. and L. meiwang Wang & Li sp. nov. but can be distinguished by median apophysis harrow-like, tip with 5 small teeth ; from L. shicheng Wang & Li sp. nov. by the tip of conductor undulate (L. shicheng Wang & Li sp. nov.); from L. zakou Wang & Li sp. nov. by the teeth of median apophysis needle-shaped in L. zakou Wang & Li sp. nov.; from L. meiwang Wang & Li sp. nov. by the tibia spines Ⅰ strongest, tip asymmetrically bifurcated (tibia spines Ⅱ strongest in L. meiwang Wang & Li sp. nov.).ll teeth  . Total length 1.63 . Similar to male in color and general features, but larger and with longer legs. Total length 2.05 (gth 2.05 -B. Carapgth 2.05 : spermatDistribution. China (Hunan).Type material. Holotype: male (IZCAS), Shicheng Cave, N27.31°, E109.07°, Jiangwu, Shanshi Town, Lianhua County, Pingxiang City, Jiangxi Province, China, 14 November 2015, Z. Chen & G. Zhou leg. Paratypes: 2 males and 5 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. sexdentataL. longyu Wang & Li sp. nov., L. zakou Wang & Li sp. nov. and L. meiwang Wang & Li sp. nov. but can be distinguished by the harrow-like median apophysis, with 10 small teeth distally ; conductor smooth ; from L. zakou Wang & Li sp. nov. by the teeth of median apophysis needle-shaped in L. zakou Wang & Li sp. nov.; from L. meiwang Wang & Li sp. nov. by the tibia spines Ⅰ strongest, tip asymmetrically bifurcated (L. meiwang Wang & Li sp. nov.).distally  . Total length 2.40 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.60 (gth 2.60 -B. Carapgth 2.60 : spermatDistribution. China (Jiangxi).Type material. Holotype: male (IZCAS), Zakou Cave, N29.35°, E109.58°, Hongyanxi Town, longshan City, Hunan Province, China, 10 January 2016, Z. Chen & Z. Wang leg. Paratypes: 3 males and 5 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. sexdentataL. longyu Wang & Li sp. nov., L. shicheng Wang & Li sp. nov., and L. meiwang Wang & Li sp. nov. but can be distinguished by on the male pedipalpal bulb median apophysis with 6 teeth, needle-shaped ; from L. shicheng Wang & Li sp. nov. by the distally undulate conductor (L. shicheng Wang & Li sp. nov.); from L. meiwang Wang & Li sp. nov. by the tibia Ⅰ spine strongest, tip asymmetrically bifurcated (L. meiwang Wang & Li sp. nov.).e-shaped  . Total length 1.75 . Similar to male in color and general features, but larger and with shorter legs. Total length 1.70 (gth 1.70 -B. Carapgth 1.70 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Meiwang Cave, N28.09°, E111.43°, Nanhua, Zhenshang Town, Lodi County, HuNan Province, China, 27 March 2016, Y. Li & Z. Chen leg. Paratypes: 1 male and 1 female, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. sexdentataL. longyu Wang & Li sp. nov., L. shicheng Wang & Li sp. nov. and L. zakou Wang & Li sp. nov. but can be distinguished by the harrow-like median apophysis, with 5 sharp teeth distally ; conductor short, reduced ; from L. zakou Wang & Li sp. nov. by the teeth of median apophysis needle-shaped in L. zakou Wang & Li sp. nov.distally , tibia Ⅱdistally  : total length 1.75 .Type material. Holotype: male (IZCAS), Xianren Cave, N29.18°, E109.95°, Xianren, Tawo Town, Yongshun County, Hunan Province, China, 14 January 2016, Z. Chen & Z. Wang leg. Paratypes: 2 males and 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. arvanitidisi Wang & Li, 2016, L. paragamiani Wang & Li, 2016 and L. erlong Wang & Li sp. nov. but can be distinguished by on the male pedipalpal bulb median apophysis with 4 teeth distally ; tibia spines Ⅰ strongest, tip asymmetrically bifurcated, spines Ⅱ, Ⅲ equally strong, stronger than other 2 ; from L. arvanitidisi by the conductor C tile-shaped (L. arvanitidisi).distally  . Total length 1.90 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.00 (gth 2.00 -B. Carapgth 2.00 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Erlong Cave, N27.82°, E110.23°, Siqian Town, Chenxi County, Huaihua City, Hunan Province, China, 19 March 2016, Y. Li & Z. Chen leg. Paratypes: 4 males and 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. arvanitidisi Wang & Li, 2016, L. paragamiani Wang & Li, 2016 and L. tawo Wang & Li sp. nov. but can be distinguished by on the male pedipalpal bulb median apophysis with 5 teeth distally ; from L. paragamiani and L. tawo Wang & Li sp. nov. by the tibia spines Ⅱ -Ⅴ slender, curved, and equally strong ; from L. arvanitidisi by the conductor C tile-shaped (L. arvanitidisi).distally  : total length 1.95 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.30 (gth 2.30 -B. Prosogth 2.30 : spermatDistribution. China (Hunan).Type material. Holotype: male (IZCAS), Wuming Cave, N25.75°, E107.92°, Dabian, Sandong Town, Sandu County, Qiannan Prefecture, Guizhou, China, 22 March 2013, H. Zhao & J. Liu leg. Paratypes: 2 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. thraciaL. chuan Wang & Li sp. nov., but can be distinguished by the male pedipalal tibia with 3 spines prolaterally, 5 slender spines, retrolaterally ; tip of median apophysis bent upwards, with 3 larger teeth distally ; conductor thin, tongue shaped (L. thracia and L. chuan Wang & Li sp. nov.).aterally -D . Total length 2.38 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.40 (gth 2.40 -B. Carapgth 2.40 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Chuan Cave, N27.08°, E105.67°, Yangchangba Town, Dafang County, Guizhou Province, China, 13 March 2011, H. Chen & Z. Zha leg. Paratype: 1 female, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. thraciaL. dabian Wang & Li sp. nov., but can be distinguished by the male pedipalpal tibia with 7 long setae prolaterally, 5 slender spines retrolaterally, with spines Ⅰ, Ⅱ, Ⅲ equally strong, stronger than others ; tip of median apophysis bent downwards, with 5 larger teeth distally ; from L. dabian Wang & Li sp. nov. by the triangular conductor (L. dabian Wang & Li sp. nov.).n others  . Total length 2.10 .Type material. Holotype: male (IZCAS), nameless Cave, N25.10°, E107.65°, Lihu Town, Nandan County, Hechi City, Guangxi Zhuang Autonomous Region, China, 31 January 2015, Y. Li & Z. Chen leg. Paratypes: 2 males and 5 females, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. notabilis , L. sexdigiti ; and L. shuang Wang & Li sp. nov., but can be separated from L. notabilis by the male pedipalpal tibia spines Ⅰ bifurcate (L. notabilis); from L. shuang Wang & Li sp. nov. by the conductor C tile-shaped, distal edge of median apophysis with 6 teeth (L. shuang Wang & Li sp. nov.); from L. sexdigiti by the strongly twisted spermathecae (spermathecae loosely twisted in L. sexdigiti).ifurcate  . Total length 2.13 . Similar to male in color and general features, but larger and with shorter legs. Total length 2.50 (gth 2.50 -B. Carapgth 2.50 : spermatDistribution. China (Guangxi).Type material. Holotype: male (IZCAS), Shuang Cave, N25.93°, E107.26°, Bailong Town, Pingtang County, Qiannan Prefecture, Guizhou Province, China, 24 July 2012, H. Zhao leg. Paratypes: 2 females, same data as holotype; 2 males and 6 females, Dongkou Cave, N25.93°, E107.25°, Longxiang, Bailong Town, Pingtang County, Qiannan Prefecture, Guizhou Province, China, 25 July 2012, H. Zhao leg.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. notabilis , L. sexdigiti , and L. lihu Wang & Li sp. nov., but can be separated from L. notabilis by the male pedipalp tibia spines Ⅰ bifurcate (L. notabilis); from L. sexdigiti and L. lihu Wang & Li sp. nov. by the conductor triangular, distal edge of median apophysis with 7 teeth (L. sexdigiti and L. lihu Wang & Li sp. nov.); from L. sexdigiti by in the female spermathecae strongly twisted (L. sexdigiti).ifurcate  . Total length 2.00 . Similar to male in color and general features, but smaller and with shorter legs. Total length 1.98 (gth 1.98 -B. Carapgth 1.98 : spermatDistribution. China (Guizhou).Type material. Holotype: male (IZCAS), Encun Cave, N25.08°, E107.59°, En, Chengguan Town, Nandan County, Hechi City, Guangxi Zhuang Autonomous Region, China, 30 January 2015, Y. Li & Z. Chen leg. Paratypes: 1 male and 1 female, same data as holotype.Etymology. The specific name refers to the type locality; noun.Diagnosis. This new species is similar to L. robustispina  but can be distinguished by the male pedipalpal tibia with 5 spines retrolaterally, with spines Ⅰ longest, spines Ⅰ, Ⅱ, Ⅲ equally strong, stronger than others (L. robustispina).n others , distal n others  . Total length 2.00 .Leptonetela zhaiMaterial examined. 4 females (IZCAS), Rudong Cave, N25.57°, E110.62°, Longpan Mountain, Dongtian, Hucheng Town, Xing'an County, Guilin City, Guangxi Zhuang Autonomous Region, China, 08 November 2012, Z. Chen & Z. Zhao leg.Description. Male. See Female. Total length 2.12 (gth 2.12 -B. Carapgth 2.12 : spermatDistribution. China (Guangxi).Remarks. The female of the species is described for the first time. Females of Leptonetelazhai were collected from the same cave where the male holotype of L. zhaiLeptoneta tianxinensis Tong & Li, 2008: 382, Type material examined. Paratypes: 12 males, 6 females (IZCAS), Tianxin Cave, N33.35°, E111.88°, Sandaohe, Qilipo Town, Neixiang County, Henan Province, China, 24 June 2005, Q. Wang & Y. Tong leg.Remarks. Our research showed that this species should be transferred to the genus Leptonetela, based on the result of DNA barcoding and morphological characters such as the pedipalpal femur lacking spines and the tibia with one strong spine retrolaterally.Guineta gigachela Lin & Li, 2010: 6, Type material examined. Holotype: male (IZCAS), Qingzi Cave, N26.51°, E107.99°, Mianxi, Sankeshu Town, Kaili City, Guizhou Province, China, 26 May 2007, Y. Li & J. Liu leg. Paratypes: 2 males and 12 females, same data as holotype.Remarks. Our studies showed that that Guineta Lin & Li, 2010 syn. nov. should be a junior synonym of Leptonetela Kratochvíl, 1978.Sinoneta notabilis Lin & Li, 2010: 83, Type material examined. Holotype: male (IZCAS), Hebiandong Cave, Kaikou Town, Duyun City, N26.00°, E107.20°, Guizhou Province, China, 8 May 2006, Y. Li leg. Paratypes: 1 male and 1 female, same data as holotype.Remarks. Our studies showed that Sinoneta Lin & Li, 2010 syn. nov. should be a junior synonym of Leptonetela Kratochvíl, 1978.Sinoneta sexdigiti Lin & Li, 2010: 87, Type material examined. Holotype: male (IZCAS), Qiaotou Cave, Dashan, Shuangliu Town, Kaiyang County, N26.05°, E107.85°, Guizhou Province, China, 11 May 2006, Y. Li & Z. Yang leg. Paratypes: 5 males and 29 females, same data as holotype.Qianleptoneta palmata Chen, Jia & Wang, 2010: 2902, Sinoneta palmataQianleptoneta).Material examined. 1 male and 1 female (IZCAS), Sanchahe Cave, N26.53°, E107.70°, Sanchahe, Jialiang Town, Libo County, Guizhou Province, China, 16 May 2011, C. Wang & L. Lin leg.Etymology. The specific name refers to the type locality; noun.Remarks. Qianleptoneta palmata was collected from Sanchahe Cave in Guizhou, China and published by Chen et al. in December 2010. Qianleptoneta palmata Chen et al, 2010 to the genus Sinoneta Lin & Li, 2010. Nevertheless, in this study our results confirmed that Qianleptoneta palmata belonged to the genus Leptonetela.Leptonetela palmata is a preoccupied name (secondary homonym) for a species collected from Dixian Cave in Guizhou, China and published by Lin & Li, in August 2010. Subsequently, Leptonetela sanchahe Wang & Li nom. nov. is proposed for the taxon from Sanchahe Cave, in Guizhou, China.Yi Wu and Guo Zheng helped to prepare photos of the manuscript. Sarah C. Crews kindly checked the English of the manuscript. Peter Jäger, Yanfeng Tong and Yucheng Lin provided valuable comments on an early version of the manuscript.