Eyewire II Proofread Cells Main List - Cell types and properties 2026-01-23.csv

Cell type name,Cell class,Short name for MS,Helmstaeder name,EW1 name,Other names,EW II Type,Reference,Reference links,temporary comments,Stratification,peak position,Axon ,Dendritic field size,Dendritic field shape,Dendrite shape ,Dendritic field density,Soma layers,Surround suppression level,Response kinetics,Response polarity,Spiking,OS,DS,Count,Example
A1,AC,A1,-,-,nNOS-1,,"Park et al., 2020",,"has been called A1 in primates, cat and rabbit ",bistratified low/deep,,yes,wide,,,,GCL,,,ON,,,,93,D234
A2,AC,A2,H24,-,,,,,,bistratified diffuse ,0,no,small ,symmetric,curly,dense,INL,,,ON,,,,83,G374
A17,AC,A17,H49,-,,,"Nelson & Kolb, 1985",,,monostratified diffuse,0,,wide,symmetric,wavy,dense,INL/GCL,,,ON,,,,79,D88
Bowtie deep in IPL,AC,Bowtie,H28,-,WF2,,"Bloomfield & Miller, 1986",,likely has subtypes ,monostratified layered,,,wide,,,,INL,,,OFF,,,,11,C482
Comet,AC,Comet,H31,-,,,,,,monostratified layered,,no,wide,asymmetric,,,INL,,,OFF,,,,2,G373
Medium-field deep in IPL sustained,AC,MfDeep-s,,-,,,,,,,,,medium ,,,,GCL,,,,,,,6,
Medium-field deep in IPL transient,AC,MfDeep-t,,-,,,,,,,,,medium ,,,,GCL,,,,,,,8,
Medium-field dorsal pointing low in IPL,AC,MfDeep-d,,-,,,,,,,,,medium ,,,,GCL,,,,,,,3,C199
Medium-field hugging ON ChAT,AC,MfHug,,-,,,,,,,,,medium ,,,,GCL,,,,,,,,C167
Medium-field low in IPL sustained,AC,MfLow-s,,-,"CRH-2, GBX2-S5",,,,,,,,medium ,,,,GCL,,,,,,,52,D308
Medium-field low in IPL transient,AC,MfLow-t,H55 ?,-,VIP3,,"Perez de Sevilla Mueller et al., 2019",,,monostratified ,1,,medium ,slightly asymmetric,branchy wavy,dense-ish,GCL,,,,,,,73,
Medium-field Mid-IPL smooth,AC,MfMid-s,,-,nGnG ? ,,,,,,,no,medium ,asymmetric,,,GCL,,,,,,,51,C334
Medium-field Mid-IPL varicose,AC,MfMid-v,H43,-,Gbx2-S3,,,,,,,no,medium ,asymmetric,,,GCL,,,,,,,29,D202
Midnight,AC,MN,,-,,,,,,,,no,wide,,,,GCL,,,ON,,,,36,D78
nNOS-2,AC,nNOS-2,"H37, H41",-,,,"Jacoby et al., 2018",,likey has sub types,monostratified deep,,,wide,,,,INL/GCL,,,ON,,,,24,
Separate dendrites and axons,AC,SAD,H57,-,,,,,at least two types,monostratified flat,0,yes,wide,asymmetric,"long curly, short cury",,GCL,,,ON,,,,144,D63
ON SAC,AC,On-SAC,H51,-,,,,,,,,no,medium ,symmetric,,,GCL,,,ON,,,,417,D363
OFF SAC,AC,Off-SAC,H33,-,,,,,,,,no,medium ,symmetric,,,INL,,,OFF,,,,113,G142
Dorsal-pointing neurites low in IPL,AC,DpLow,,-,,,,,,,,,,,,,,,,,,,,2,
Wide-field deep in IPL,AC,WfDeep,,-,,,,,,,,,wide,,,,,,,,,,,,
Wide-field mid IPL,AC,WfMid,H37,-,,,,,,,,,wide,,,,,,,,,,,59,D223
Wide-field ON non-spiny dense,AC,WfNs-d,H56,-,,,,,,monostratified flat,0,,wide,asymmetric,straight,dense,GCL,,,ON,,,,39,D273
Wide-field ON non-spiny sparse,AC,WfNs-s,displaced H53,-,,,,,,,,,wide,,,,,,,ON,,,,21,D44
Wide-field ON spiny spikelets,AC,WfS-sl,,-,,,,,,,,probably,wide,,,,,,,ON,,,,31,D104
Wide-field ON spiny spikes,AC,WfS-s,,-,,,,,,,,probably,wide,,,,,,,ON,,,,28,D342
Bursty suppressed by contrast,RGC,bSbC,,2o,,,"Tien et al., 2016 ?","Jacoby et al., 2018",,,,,,,,,GCL,,,,,,,2,D203
M1,RGC,M1,,1ws,,,,,,,,,,,,,GCL,,,,,,,4,D366
F-mini-OFF,RGC,FmOff,,2an,,,"Rousso et al., 2016","Nath et al., 2016",,,,,small ,,,,GCL,,,OFF,,yes,yes ,10,D49
F-mini-ON,RGC,FmOn,,63,,,"Rousso et al., 2016","Nath et al., 2016",,,,,small ,,,,GCL,,,ON,,yes,yes ,9,D69
HD1,RGC,HD1,,5si,,,"Jacoby et al., 2017",,,,,,small ,,,,GCL,,transient ,ON-OFF,,,,5,D311
HD2,RGC,HD2,,5so,"W3b, Object Motion Sensor",,"Zhang et al., 2012","Jacoby et al., 2017",,,,,small ,,,,GCL,strong,transient ,ON-OFF,,,,7,D45
Local edge detector,RGC,LED,H7,51,,,,,,,,,,,,,GCL,,,,,,,,E8
M2,RGC,M2,,9w,,,,,,,,,,,,,GCL,,,,,,,,
M6,RGC,M6,,91,,,,,,,,,,,,,GCL,,,,,,,4,D180
5to,RGC,e5to,,5to,,,,,,,,,,,,,GCL,,,,,,,,
OFF horizontal OS - symmetric,RGC,hOffOS,,2aw,F-midi-OFF,,,,,,,,,,,,GCL,,,OFF,,yes,,,D324
OFF medium sustained,RGC,sOff-m,,2i,,,,,,,,,,,,,GCL,,sustained,OFF,,,,4,D138
OFF OS - subtype unknown,RGC,OffOS,,-,,,,,,,,,,,,,GCL,,,OFF,,yes,,,
OFF sustained alpha,RGC,sOff-a,,1wt,,,,,,,,,,,,,GCL,,sustained,OFF,,,,1,D271
OFF sustained EW1no,RGC,e1no,,1no,,,,,,,,,,,,,GCL,,sustained,OFF,,,,1,D235
OFF sustained EW3o,RGC,e3o,,3o,,,,,,,,,,,,,GCL,,sustained,OFF,,,,1,
OFF transient - subtype unknown,RGC,tOff,,-,,,,,,,,,,,,,GCL,,transient ,OFF,,,,,
OFF transient alpha,RGC,tOff-a,,4ow,,,,,,,,,,,,,GCL,,transient ,OFF,,,,4,D275
OFF transient medium RF,RGC,tOff-m,,4on,,,,,,,,,,,,,GCL,,transient ,OFF,,,,4,D20
OFF transient small RF,RGC,tOff-s,,4i,,,,,,,,,,,,,GCL,,transient ,OFF,,,,5,D357
OFF vertical OS - asymmetric,RGC,vOffOS,,2aw,"JamB, OFF DS",,"Kim et al., 2008",,,,,,,,,,GCL,,transient ,OFF,,yes,yes ,14,D14
ON alpha,RGC,sOn-a,,8w,M4,,,,,,,,,,,,GCL,,,ON,,,,7,D39
ON bursty,RGC,bOnSbC,,3i,,,,,,,,,,,,,GCL,,,ON,,,,3,D189
ON delayed,RGC,dOnSbC,,73,transient SbC,,"Jacoby et al., 2016 ?","Jacoby et al., 2018",,,,,,,,,GCL,,,ON,,,,4,D5
ON DS sustained - direction unknown,RGC,sOnDS,,"7iv,r,d",,,,,,,,,,,,,GCL,,sustained,ON,,,yes ,2,D34
ON DS sustained - dorsonasal,RGC,sOnDS-dn,,7iv,,,,,,,,,,,,,GCL,,sustained,ON,,,yes ,1,
ON DS sustained - temporal,RGC,sOnDS-t,,7ir,,,,,,,,,,,,,GCL,,sustained,ON,,,yes ,2,
ON DS sustained - ventral,RGC,sOnDS-v,,7id,,,,,,,,,,,,,GCL,,transient ,ON,,,yes ,1,
ON DS transient - temporal,RGC,tOnDS-t,,7o,,,,,,,,,,,,,GCL,,,ON,,,yes ,,
ON OS large RF - orientation unknown,RGC,OnOS-l,,-,,,,,,,,,,,,,GCL,,,ON,,yes,,,
ON OS small RF - orientation unknown,RGC,OnOS-s,,-,,,,,,,,,,,,,GCL,,,ON,,yes,,,
ON small OFF large,RGC,On-s-Off-l,,1ni,,,,,,,,,,,,,GCL,,,ON,,,,,E23
ON transient EW6t,RGC,e6t,,6t,,,,,,,,,,,,,GCL,,transient ,ON,,,,2,D19
ON transient medium RF,RGC,tOn-m,,6sw,,,,,,,,,,,,,GCL,,transient ,ON,,,,3,D9
ON transient small RF,RGC,tOn-s,,6sn,,,,,,,,,,,,,GCL,,transient ,ON,,,,3,D113
ON vertical OS large RF,RGC,vOnOS-l,,tbd,,,"Nath et al., 2016",,,,,,,,,,GCL,,,ON,,yes,,,
ON vertical OS small RF,RGC,vOnOS-s,,tbd,,,"Nath et al., 2016",,,,,,,,,,GCL,,,ON,,yes,,1,
ON horizontal OS small RF,RGC,hOnOS-s,,tbd,,,,,,,,,,,,,GCL,,,ON,,yes,,1,
ON horizontal OS large RF,RGC,hOnOSl,,tbd,,,,,,,,,,,,,GCL,,,ON,,yes,,,
ON-OFF - subtype unknown,RGC,OO,,-,,,,,,,,,,,,,GCL,,,ON-OFF,,,,7,
ON-OFF DS - direction unknown,RGC,OODS,H9,37,,,,,,,,,,,,,GCL,,,ON-OFF,,,,8,
ON-OFF DS - dorsal,RGC,OODS-d,,37v,,,,,,,,,,,,,GCL,,,ON-OFF,,,,,
ON-OFF DS - nasal,RGC,OODS-n,,37c,,,,,,,,,,,,,GCL,,,ON-OFF,,,,,
ON-OFF DS - temporal,RGC,OODS-t,,37r,,,,,,,,,,,,,GCL,,,ON-OFF,,,,,
ON-OFF DS - ventral,RGC,OODS-v,,37d,,,,,,,,,,,,,GCL,,,ON-OFF,,,,1,D2
PixON,RGC,M5,,9n,M5,,,,,,,,,,,,GCL,,,,,,,5,E103
Sustained suppressed-by-contrast no surround EW27,RGC,e27,,27,,,"Goetz et al., 2022",,,,,,,,,,GCL,,,,,,,1,D27
Sustained suppressed-by-contrast strong surround EW28,RGC,e28,,28,,,"Goetz et al., 2022",,,,,,,,,,GCL,,,,,,,4,D157
UHD,RGC,UHD,,5ti,W3b,,"Jacoby et al., 2017",,,,,,small ,,,,GCL,,transient ,ON-OFF,,,,16,D42
EW25,RGC,e25,,25,,,,,,,,,,,,,GCL,,,,,,,8,D95
ON sustained - subtype unknown,RGC,sOn,,1ws,,,,,,,,,,,,,GCL,,,,,,,,D181
EW915,RGC,e915,,915,,,,,,,,,,,,,GCL,,,,,,,,
EW85,RGC,e85,,85,,,,,,,,,,,,,GCL,,,,,,,,
H13,AC,H13,H13,,,,,,,,,,small ,,,,INL,,,OFF,,,,25,
H14,AC,H14,H14,,Narrow S1,,MacNeil & Masland 1999,,,,,,small ,,,,INL,,,OFF,,,,9,
H15,AC,H15,H15,,Flat bistratified,,MacNeil & Masland 1999,,,,,,small ,,,,INL,,,OFF,,,,7,
H16,AC,H16,H16,,,,,,,,,,small ,,,,INL,,,ON-OFF,,,,3,
H18,AC,H18,H18,,A8 ,,"Lee et al., 2016","Jo et al., 2023 say SEG looks like H18. Their SEG looks different from the one in Kay et al., 2011",,,,,small ,,,,INL,,,ON-OFF,,,,30,G199
H19,AC,H19,H19,,OFF NAC,,"Jain et al, 2022",,,,,,small ,,,,INL,,,OFF,,,,34,
H20,AC,H20,H20,,"SEG, Narrow diffuse, nGnG",,"Kay et al., 2011",,,,,,small ,,,,INL,,,ON-OFF,,,,46,G887
H21,AC,H21,H21,,Flag A,,MacNeil & Masland 1999,,,,,,small ,,,,INL,,,ON-OFF,,,,17,G1049
H22,AC,H22,H22,,"ON NAC, SI - AC",,"Jo et al., 2023",,,,,,small ,,,,INL,,,,,,,3,
H23,AC,H23,H23,,"MAC, SI - AC?",,"Grimes et al., 2021",,,,,,small ,,,,INL,,,ON,,,,56,
vGlut3,AC,vGlut3,"H46, H48 ? ",,GAC,,"Lee et al., 2016","Mani et al., 2023",,diffuse between ChAT bands,,,,,,,INL,,transient ,ON-OFF,,,,,
VIP1,AC,VIP1,H40,,VIPm-RAC2,,"Perez de Sevilla Mueller et al., 2019",Zhu et al;. 2014,,,,,,,,,,,,,,,,,
VIP2,AC,VIP2,H50,,,non-displaced hugging ON ChAT,"Perez de Sevilla Mueller et al., 2019",,small ?,monostratified diffuse,2,,medium ,symmetric,curly,dense,,,,,,,,,
DAC ,AC,DAC ,H25,,dopaminergic amacrine cells,,"Zhang et al., 2008",,,,,,,,,,,,,,,,,,
t1,BC,t1,,,,,"Shekar et al., 2017",,,,,,,,,,INL,,transient ,OFF,,,,38,G13
t2,BC,t2,,,,,,,,,,,,,,,INL,,transient ,OFF,,,,86,G14
t3a,BC,t3a,,,,,,,,,,,,,,,INL,,transient ,OFF,,,,24,G23
t3b,BC,t3b,,,,,,,,,,,,,,,INL,,transient ,OFF,,,,31,G25
t4,BC,t4,,,,,,,,,,,,,,,INL,,transient ,OFF,,,,47,G29
t5t,BC,t5t,,,,,,,,,,,,,,,INL,,sustained,ON,,,,96,G51
t5i,BC,t5i,,,,,,,,,,,,,,,INL,,sustained,ON,,,,38,
t5o,BC,t5o,,,,,,,,,,,,,,,INL,,sustained,ON,,,,96,
XBC,BC,XBC,,,,,,,,,,,,,,,INL,,sustained,ON,,,,18,G62
t6,BC,t6,,,,,,,,,,,,,,,INL,,sustained,ON,,,,133,G63
t7,BC,t7,,,,,,,,,,,,,,,INL,,sustained,ON,,,,67,G79
t8,BC,t8,,,,,,,,,,,,,,,INL,,sustained,ON,,,,20,G82
t9,BC,t9,,,,,,,,,,,,,,,INL,,sustained,ON,,,,12,G84
RBC,BC,RBC,,,,,,,,,,,,,,,INL,,sustained,ON,,,,158,G93
NBC,BC,NBC,,,Non-canonical OFF strat mGluR6+ ON BC,,,,,,,,,,,,INL,,,,,,,6,
GluMi,BC,GluMi,,,,,"Santina et al., 2016",,,,,,,,,,INL,,,,,,,,
Horizontal cell,HC,HC,,,,,,,,,,,,,,,INL,,,,,,,,
Müller cell,Glia,Müller,,,,,,,,,,,,,,,,,,,,,,,
Astrocyte,Glia,Astro,,,,,,,,,,,,,,,,,,,,,,,
Microglia,Glia,mGlia,,,,,,,,,,,,,,,,,,,,,,,
Endothelial cell ,Vascular,Endo,,,,,,,,,,,,,,,,,,,,,,,
Pericyte,Vascular,Peri,,,,,,,,,,,,,,,,,,,,,,,
,,,,,,,,,,,,,,,,,,,,,,,,,
"Helmstaedter AC types (grey is assigned to type on the lis, white to be added)",,,,,,,,,,,,,,,,,,,,,,,,,
H24,,,,,A2,,,,,,,,,,,,,,,,,,,,
A8,,,H42? or H18/H20,,,,"Lee et al., 2016",,,,,,medium ,,,,INL,,,ON-OFF,,,,,
H25,,,DAC,,WF1,,MacNeil & Masland 1999,,,,,,medium ,,,,INL,,,,,,,,
H26,,,,,,,,,,,,,medium ,,,,INL,,,OFF,,,,,
H27,,,,,,,,,,,,,medium ,,,,GCL,,,OFF,,,,,
H28,,,,,,non-dosplaced bowtie ,,,,,,,wide,elongated,straight,sparse,INL,,,,,,,,
H29,,,,,,,,,,,,,medium ,,,,INL,,,OFF,,,,,
H30,,,,,,,,,,,,,medium ,,,,INL,,,OFF,,,,,
H31,,,,,WF1,,,,,,,,,,,,,,,,,,,,
H32,,,,,,monostratified layered,7,no,medium,symmetric,straight/ knotty,dense ,,,,,,,,,,,,,
H33,,,,,OFF SAC,,,,,,,,,,,,,,,,,,,,
H34,,,,,interplexiform ?,,,,,,,,medium ,,,,INL,,,ON-OFF,,,,,
H35,,,,,,,,,,diffuse,,,medium ,,,,INL,,,ON-OFF,,,,,
H36,,,,,,,,,,tristratified,,,medium ,,,,INL,,,ON-OFF,,,,,
H37,,,,,,Wide-field mid IPL or nNOS2,,,,,,,,,,,,,,,,,,,
H38,,,,,,,MacNeil & Masland 1999,,,Diffuse multistratified,,,medium ,,,,INL,,,ON-OFF,,,,,
H39,,,,,WF3,,MacNeil & Masland 1999,,,,,,medium ,,,,INL,,,ON,,,,,
H40,,,,,VIP1/ VIPm,,,,,bistratified,,,medium ,,,,INL,,,ON,,,,,
H41,,,,,,Wide-field mid IPL or nNOS2,,,,,,,medium ,,,,INL,,,,,,,,
H42,,,,,,,,,,,,,,,,,INL,,,,,,,,
H43,,,,,,MF mid-IPL smooth ,,,,,,,medium ,,,,GCL,,,ON,,,,,
H44,,,,,"Fountain, bigger H18 ",,MacNeil & Masland 1999,,,,,,medium ,,,,INL,,,ON-OFF,,,,,
H45,,,,,,,,,,bistratifed diffuse,1,,wide,symmetric,wavy,"dense in center, outer sparse",INL,,,,,,,,one example found
H46,,,,,"vGlut3, Narrow s3",non-dosplaced MF mid-IPL varicose,MacNeil & Masland 1999,,,,,,medium ,,,,INL,,,ON,,,,,
H47,,,,,,non-displaced nNOS-2 ,,,,monostratified layered,3,no,wide,elongated,straight,sparse,INL,,,ON,,,,,
H48,,,,,"vGlut3, AB broad diffuse",,MacNeil & Masland 1999,,,,,,medium ,,,,INL,,,ON,,,,,
H49,,,,,A17,,,,,,,,,,,,,,,,,,,,
H50,,,,,VIP2,non-displaced hugging ON ChAT,,,,monostratified layered,2,unclear,medium ,,,,INL,,,ON,,,,,
H51,,,,,ON SAC,,,,,,,,,,,,,,,,,,,,
H52,,,,,,,,,,bistratified,,,medium ,,,,INL,,,ON,,,,,
H53,,,,,,non-displaced non-spinky sparse stratifying,,,,monostratified,2,unclear,wide,unclear,long straight,sparse,INL,,,ON,,,,,
H54,,,,,WF4,non-displaced SAD stratifying,MacNeil & Masland 1999,,,monostratified,1,yes,wide,asymmetric,"many short branchy, few long straight","dense in center, outer sparce",INL,,,,,,,,
H55,,,,,MF low in IPL transient/sustained,,,,weakly bistatified,,,,medium ,,,,GCL,,,ON,,,,,
H56,,,,,,Wide-field ON non-spiny dense,,,,,,,medium ,,,,GCL,,,ON,,,,,
H57,,,,,,SAD,,,,,,,medium ,,,,GCL,,,ON,,,,,
,,,,,,,,,,,,,,,,,,,,,,,,,
Other Types  (to be added),,,,,,,,,,,,,,,,,,,,,,,,,
VIPm-RAC2,,,H40,,,,from David,,,bistratified,,,,,,,,,,,,,,,
RAC2,,,,,"just a few branches that go into the inner ON, otherwise very similar to VIPm-RAC2",,from David,,,,,,,,,,,,,,,,,,
nGnG,,,,,,,"Kay et al., 2011",,,,,,,,,,,,,,,,,,
WA1-5,,,,,,,Lin & Masland 2006,,,,,,,,,,,,,,,,,,
B/K AC (and subtypes),,,,,,,"Park et al., 2024",,,,,,,,,,,,,,,,,,
bifid/waterfall/.. ; cluster1-8,,,,,,,"Badea & Nathans, 2004",,,,,,,,,,,,,,,,,,
interplexiform,,,H34,,,,"Dedek et al., 2009",,,,,,,,,,,,,,,,,,
CA1,,,,,,,"Yan et al., 2020",,,,,,,,,,,,,,,,,,
CA2,,,,,,,"Yan et al., 2020",,,,,,,,,,,,,,,,,,
polyaxonal ?,,,,,"SAD, A1 ",,"Voegly et al, 2001",,,,,,,,,,,,,,,,,,
VG3,,,,,,,"Yan et al., 2020",,,,,,,,,,,,,,,,,,
VIP3,,,,,MF low in IPL transient ?,,"Perez de Sevilla Mueller et al., 2019",,,,,,,,,,,,,,,,,,
,,,,,,,,,,,,,,,,,,,,,,,,,
,,,,,,,,,,,,,,,,,,,,,,,,,
NEW EW II TYPES,,,,,,,,,,,,,,,,,,,,,,,,,
SAD medium,,,,,,,,,,,,,,,,,,,,,,,,,
SAD large,,,,,,,,,,,,,,,,,,,,,,,,,
SAD giant,,,,,,,,,,,,,,,,,,,,,,,,,
deeply SAD,,,,,,,,,,,,,,,,,,,,,,,,,
non-displaced MF low in IPL ,,,,,,,,,,monostratified diffuse,2,no,medium,symmetric,wavy,dense,INL,,,,,,,,
non-displaced nNOS-2,,,,,,,,,,,,,,,,,,,,,,,,,
H32,,,,,,non-displaced MF deep in IPL ,,,,monostratified layered,7,no,medium,symmetric,straight/ knotty,dense ,,,,,,,,,
nNos-1,,,,,,,,,,,,,,,,,,,,,,,,,
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example found ,,,,,,,,,,,,,,,,,,,,,,,,,
new match for existing types (literature or Julia),,,,,,,,,,,,,,,,,,,,,,,,,
confirmed match,,,,,,,,,,,,,,,,,,,,,,,,,
new type ,,,,,,,,,,,,,,,,,,,,,,,,,